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Kinda boils down to film study and opponent tendencies. If through film study you find out that the opponent's offense throws "quick game" on downs that are 3rd & medium, it may not make much sense to send 4 (or more) rushers because you know there's an 85% chance the ball is coming out of the...

Because if you're expecting "quick game" (QB catch and throw) it makes no sense to send 4 rushers. Some DC's feel it's better to use that 4th rusher to sit underneath a quick hitch (or any other quick route at the sticks).